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Taxa hierarchy  

family Capniidae Banks, 1900

• Common name(s):  Snowflies
• Distribution:
  •  
• Ecology:
  • Freshwater, terrestrial. Shredders of decayed tree leaves
• Citations (38):
  • Banks. 1900[1899]. Trans. Amer. Ent. Soc. 26(3):239-259  
  • Klapálek, Frantisek. 1905. Cas. ces. Spol. ent. 2:27-32 >> Note: New family.
  • Klapálek, Frantisek. 1909. In Klapálek, Frantisek & Grunberg. Hft. 8. Ephemerida, Plecoptera, Lepidoptera. In Brauer, A. Die Süsswasserfauna Deutschlands. Eine Exkursionsfauna 55  
  • Tillyard. 1921. Canad. Ent. 53:40  
  • Frison. 1935. Ill. Nat. Hist. Surv. Bull. 20:310, 355  
  • Illies. 1966. Das Tierreich 82:121 >> Note: Capniidae
  • Raušer. 1968. Entomologische Abhandlungen 34(5):337-343
  • Harper, P.P. & Hynes. 1971. Can. J. Zool. 49:921-940 >> Note: Keys to males, females and nymphs of eastern Canada.
  • Zwick, P. 1973. Das Tierreich 94:359 >> Note: Capniidae
  • Zhiltzova & Levanidova. 1978. Trud. zool. Inst. Akad. Nauk 61:3
  • Donald & Patriquin. 1983. Canad. Ent. 115:921-926
  • Zwick, P. 1990. In Campbell [Ed.]. Mayflies and Stoneflies: Life Histories and Biology. Proceedings of the 5th International Ephemeroptera Conference and the 9th International Plecoptera Conference. Ser. Entomol. 44:141-148
  • Mangum. 1999. Journal of Freshwater Ecology 14(1):129-130  
  • Zwick, P. 2000. Annu.Rev.Entomol. 45:722
  • Nelson, C.R. & Giuliani. 2001. In Dominguez [Ed.]. Trends in Research in Ephemeroptera and Plecoptera. 199-208
  • Zhiltzova. 2001. In Dominguez [Ed.]. Trends in Research in Ephemeroptera and Plecoptera. 423-430
  • Stewart & Stark. 2002. Caddis Press 113
  • Zhiltzova. 2003. Fauna of Russia and Neighbouring Countries New Series No. 145:316
  • Stewart & Sandberg. 2006. In Drosopoulos & Claridge. Insect Sounds and Communication: Physiology, Behaviour, Ecology and Evolution 184, 185
  • Zwick, P. 2006. Aquatic Insects 28(1):13-22
  • Nations, Stark & Hicks. 2007. Illiesia 3(9):70-94  
  • Sheldon. 2008. In Hauer, Stanford & Newell [Ed.]. International Advances in the Ecology, Zoogeography, and Systematics of Mayflies and Stoneflies 15-38  
  • Alibozek & Ganger. 2008. Journal of Freshwater Ecology 23(4):673  
  • Eriksen, Amekleiv & Kjaerstad. 2009. Journal of Freshwater Ecology 24(4):597  
  • Heinold. 2010. Master's Thesis 92
  • Menninger. 2011. Scientific American Blog 2/24/2011  
  • Judson & C.R. Nelson. 2012. Zootaxa 3541:17
  • Murányi, Gamboa & Orci. 2014. Zootaxa 3812:82 pp.  
  • Murányi, W-H. Li & D. Yang. 2015. Zootaxa 4059(2):371-382  
  • Anzani, Krisanti & Wardiatno. 2016. Journal of Biodiversity and Environmental Sciences 9(1):128
  • Yang, D. & W-H. Li. 2018. Species Catalogue of China. Vol. 2. Animals, Insecta (III), Plecoptera 1 >> Note: Capniidae.
  • Bogan & S. M. Carlson. 2018. Illiesia 14(8):144-154  
  • Hohmann & Kuttner. 2018. Veröffentlichungen des Naturkundemuseums Potsdam 4:31
  • Broome, Stark & Baumann. 2019. Illiesia 15(1):1-26 >> Note: Reviewed Bolshecapnia, split from it two genera, Eurekapnia, Sasquacapnia.
  • Cheney, A. H. Roy, R. F. Smith & DeWalt. 2019. Environmental Entomology 20(20):1-11
  • Chen, Z-T. 2019. Zoosymposia 16:64
  • Burton. 2019. Canadian Journal of Arthropod Identification 36:5 >> Note: Key to the adult Capniidae of Canada east of Alberta.
  • Dénes, Murányi, Soós, Szocs & Keresztes. 2021. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 64(2):67 >> Note: Key to adult males of Capniidae from Romania.
• Type genus:  Capnia Pictet, F.J., 1841; priority for family-group names based on Capnia dates from Capnini Banks, 1900.  First use as Capniidae by Klapálek, Frantisek, 1905.

• synonym Apteroperlidae Matsumura, 1931
• synonym Capniinae Banks, 1900
• synonym Capnini Banks, 1900
• subfamily Capniidae
• genus Allocapnia Claassen, 1928
• synonym Capnella Claassen, 1924
• synonym Capnellula Strand, 1935
• species group forbesi Frison, 1929
• species group granulata (Claassen, 1924)
• species group illinoensis Frison, 1935
• species group jeanae Ross, 1964
• species group minima (Barnston, 1848)
• species group mystica Frison, 1929
• species group pygmaea (Burmeister, 1839)
• species group recta (Claassen, 1924)
• species group rickeri Frison, 1942
• species group virginiana Frison, 1942
• species group vivipara (Claassen, 1924)
• species oribata Poulton & Stewart, 1987
• species simmonsi Kondratieff & Voshell, 1981
• genus Apteroperla Matsumura, 1931
• synonym Allocapniella Kawai, 1955
• species babaensis (Kawai, 1967)
• species elongata (Kawai, 1967)
• species monticola (Kawai, 1955)
• species tikumana (Uéno, 1938)
• species verdea (Kawai, 1967)
• species yazawai Matsumura, 1931
• genus Arsapnia Banks, 1897
• species coyote (Nelson & Baumann, 1987)
• species decepta Banks, 1897
• species pileata (Jewett, 1966)
• species sequoia (Nelson & Baumann, 1987)
• species teresa (Claassen, 1924)
• species tumida (Claassen, 1924)
• species utahensis (Gaufin & Jewett, 1962)
• genus Baikaloperla Zapekina-Dulkeit & Zhiltzova, 1973
• species elongata Zapekina-Dulkeit & Zhiltzova, 1973
• species kozhovi Zapekina-Dulkeit & Zhiltzova, 1973
• genus Bolschecapnia Ricker, 1965
• species gregsoni (Ricker, 1965)
• species milami (Nebeker & Gaufin, 1967)
• species rogozera (Ricker, 1965)
• species spenceri (Ricker, 1965)
• genus Capnia Pictet, 1841
• nomen dubium necydaloides Pictet, 1841
• species group atra Morton, 1896
• species group barberi Nelson & Baumann, 1989
• species group bifrons Zhiltzova, 2001
• species group californica Claassen, 1924
• species group coloradensis Claassen, 1937
• species group cordata Kimmins, 1947
• species group decepta Nelson & Baumann, 1989
• species group excavata Claassen, 1924
• species group gracilaria Claassen, 1924
• species group japonica Okamoto, 1922
• species group mariposa Nelson & Baumann, 1987
• species group nana Claassen, 1924
• species group nearctica Banks, 1919
• species group pedestris Kimmins, 1947
• species group vernalis Newport, 1848
• species group vidua Klapálek, 1904
• species affinis Morton, 1896
• species asakawaena Kohno, 1952
• species bicornata Alouf, 1992
• species bifida Jewett, 1960
• species bituberculata Uéno, 1929
• species caryi Baumann & Jacobi, 2002
• species erecta Jewett, 1955
• species femina Kawai, 1968
• species fialai Nelson & Baumann, 1990
• species fukushimana Kohno, 1952
• species gibbera Jewett, 1960
• species kersti Nelson, 2004
• species kibuneana Kawai, 1957
• species kolymensis Zhiltzova, 1981
• species lepnevae Zapekina-Dulkeit, 1957
• species levanidovae Kawai, 1969
• species mitsuseana Hanada, 2018
• species naebensis Kawai, 1967
• species naraiensis Kawai, 1957
• species nelsoni Kondratieff & Baumann, 2002
• species noshaqensis Kawai, 1966
• species qilianshana Li & Yang, 2009
• species rara Zapekina-Dulkeit, 1970
• species scobina Jewett, 1966
• species sextuberculata Jewett, 1954
• species shirahatae Kohno, 1952
• species spinulosa Claassen, 1937
• species storkani Šámal, 1935
• species takahashii Okamoto, 1922
• species triangulipennis Jewett, 1975
• species tshukotica Zhiltzova & Levanidova, 1978
• species yunnana Li & Yang, 2011
• species zijinshana Du & Chen, 2016
• species zukeli Hanson, 1943
• genus Capniella Klapálek, 1920
• synonym Neocapniella Claassen, 1936
• species endemica (Zapekina-Dulkeit, 1955)
• species ghilarovi Zhiltzova, 1988
• species gibba Hwang, Li & Murányi, 2021
• species nodosa Klapálek, 1920
• genus Capnioneura Ris, 1905
• synonym Napcia Navás, 1917
• species aptera Berthélemy, 1969
• species atlasica Vinçon, El Alami & Errochdi, 2014
• species balkanica Baumann & Kaćanski, 1975
• species bolkari Vinçon & Sivec, 2011
• species brachyptera Despax, 1932
• species caucasica Zhiltzova, 1964
• species gelesae Berthélemy & Baena-Ruiz, 1984
• species gouanerae Vinçon & Sivec, 2011
• species libera (Navás, 1909)
• species mitis Despax, 1932
• species narcea Vinçon & Sánchez-Ortega, 2002
• species nemuroides Ris, 1905
• species petitpierreae Aubert, 1960
• species petricola Giudicelli, 1967
• species valandovi Ikonomov, 1978
• genus Capnopsis Morton, 1896
• synonym Capnodes Rostock, 1892
• species schilleri (Rostock, 1892)
• genus Capnura Banks, 1900
• species anas Nelson & Baumann, 1987
• species elevata (Frison, 1942)
• species fibula (Claassen, 1924)
• species intermontana Nelson & Baumann, 1987
• species manitoba (Claassen, 1924)
• species venosa Banks, 1900
• species wanica (Frison, 1944)
• genus †Dobbertiniopteryx Ansorge, 1993
• species †capniomimus Ansorge, 1993
• species †juracapnia Liu, Sinitshenkova & Ren, 2009
• genus Eocapnia Kawai, 1955
• species nivalis (Uéno, 1929)
• species shigaensis Kawai, 1967
• species yezoensis Kawai, 1955
• genus Eucapnopsis Okamoto, 1922
• species brevicauda (Claassen, 1924)
• species quattuorsegmentata Okamoto, 1922
• species stigmatica Okamoto, 1922
• species yunnana Cao, Yang & Li, 2020
• genus Eurekapnia Stark & Broome, 2019
• species maculata (Jewett, 1954)
• genus Isocapnia Banks, 1938
• species abbreviata Frison, 1942
• species agassizi Ricker, 1943
• species anguis Chen & Du, 2017
• species apicispina Chen & Du, 2017
• species aptera Zhiltzova, 1969
• species arcuata Zhiltzova, 1975
• species crinita (Needham & Claassen, 1925)
• species eichlini Zenger & Baumann, 2004
• species grandis (Banks, 1907)
• species guentheri (Joost, 1970)
• species hyalita Ricker, 1959
• species integra Hanson, 1943
• species japonica Kohno, 1953
• species kudia Ricker, 1959
• species mogila Ricker, 1959
• species orientalis Zhiltzova, 1975
• species palousa Zenger & Baumann, 2004
• species rickeri Zenger & Baumann, 2004
• species sibirica (Zapekina-Dulkeit, 1955)
• species spenceri Ricker, 1943
• species vedderensis (Ricker, 1943)
• genus Mesocapnia Raušer, 1968
• species altaica (Zapekina-Dulkeit, 1955)
• species aptera Lee & Baumann, 2011
• species arizonensis (Baumann & Gaufin, 1969)
• species autumna (Baumann & Gaufin, 1970)
• species bakeri (Banks, 1918)
• species bergi (Ricker, 1965)
• species bulbosa Nelson & Baumann, 1990
• species daxingana Chen & Du, 2016
• species frisoni (Baumann & Gaufin, 1970)
• species lapwae (Baumann & Gaufin, 1970)
• species oenone (Neave, 1929)
• species ogotoruka (Jewett, 1964)
• species porrecta (Jewett, 1954)
• species projecta (Frison, 1937)
• species silvatica Raušer, 1968
• species sugluka (Ricker, 1965)
• species variabilis (Klapálek, 1920)
• species werneri (Baumann & Gaufin, 1970)
• species yoloensis (Baumann & Gaufin, 1970)
• genus Nemocapnia Banks, 1938
• species carolina Banks, 1938
• species japonica Kohno, 1953
• genus Paracapnia Hanson, 1946
• species angulata Hanson, 1961
• species baumanni Kondratieff & Lee, 2010
• species boris Stark & Baumann, 2004
• species disala (Jewett, 1962)
• species ensicala (Jewett, 1962)
• species humboldta Baumann & Lee, 2007
• species khorensis Zhiltzova, 1972
• species leisteri Potikha & Zhiltzova, 2005
• species opis (Newman, 1839)
• species recta Zhiltzova, 1984
• species sikhotensis Zhiltzova, 1978
• genus †Rovnocapnia Sinitshenkova, 2009
• species †ambita Sinitshenkova, 2009
• species †atra Sinitshenkova, 2009
• genus Sasquacapnia Baumann & Broome, 2019
• species missiona (Baumann & Potter, 2007)
• species sasquatchi (Ricker, 1965)
• genus Sierracapnia Bottorff & Baumann, 2015
• species barberi (Claassen, 1924)
• species hornigi (Baumann & Sheldon, 1984)
• species mono (Nelson & Baumann, 1987)
• species palomar (Nelson & Baumann, 1987)
• species shepardi (Nelson & Baumann, 1987)
• species washoe Bottorff & Baumann, 2015
• species yosemite (Nelson & Baumann, 1987)
• genus Sinocapnia Murányi, Li & Yang, 2015
• species kuankuoshui Murányi, Li & Yang, 2015
• genus Takagripopteryx Okamoto, 1922
• synonym Allocapnopsis Kawai, 1955
• nomen nudum kawaii Kohno, 1964
• species imamurai Kohno, 1954
• species jezoensis Kohno, 1954
• species nigra Okamoto, 1922
• species zhuikovae Zhiltzova, 1980
• genus Utacapnia Nebeker & Gaufin, 1967
• species columbiana (Claassen, 1924)
• species distincta (Frison, 1937)
• species imbera (Nebeker & Gaufin, 1965)
• species labradora (Ricker, 1954)
• species lemoniana (Nebeker & Gaufin, 1965)
• species logana (Nebeker & Gaufin, 1965)
• species nedia (Nebeker & Gaufin, 1966)
• species poda (Nebeker & Gaufin, 1965)
• species sierra (Nebeker & Gaufin, 1965)
• species tahoensis (Nebeker & Gaufin, 1965)
• species trava (Nebeker & Gaufin, 1965)
• genus Zwicknia Murányi, 2014
• species acuta Murányi & Orci, 2014
• species avilensis Reding, Pardo & Vinçon, 2021
• species bifrons (Newman, 1838)
• species gattolliati Vinçon & Reding, 2018
• species komica Murányi & Boumans, 2014
• species kovacsi Murányi & Gamboa, 2014
• species ledoarei Reding, Launay, Ruffoni, Vinçon & Boumans, 2016
• species rupprechti Murányi, Orci & Gamboa, 2014
• species sevanica (Zhiltzova, 1964)
• species tuberculata (Zhiltzova, 1964)
• species turkestanica (Kimmins, 1950)
• species westermanni Boumans & Murányi, 2014

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