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Taxa hierarchy  

genus Neoperla Needham, 1905

• Nomenclatural details:   Nomen novum (Junior homonym is Pseudoperla)
• Images:
• Distribution:
  •  
   Eastern North America, Tropical and Temperate Asia, central Africa.
• Ecology:
  • Freshwater, terrestrial. Predators
• Specimen records are available.
• Citations (44):
  • Needham. 1905. Proc.Biol.Soc.Wash. 18:108   >> Neoperla    urn:lsid:Plecoptera.speciesfile.org:TaxonName:2854
  • Banks. 1906. Canad. Ent. 38:221-224   >> Neoperla
  • Klapálek, Frantisek. 1907. Rozpr. Ceské. Akad. 16(31):24   >> Neoperla
  • Klapálek, Frantisek. 1909. Wien. Ent. Ztg. 28(7/8):216   >> Neoperla
  • Needham & Claassen. 1925. The Thomas Say Foundation II:133, 287   >> Neoperla
  • Frison. 1935. Ill. Nat. Hist. Surv. Bull. 20:381   >> Neoperla
  • Illies. 1966. Das Tierreich 82:259, 261, 267 >> Neoperla
  • Zwick, P. 1973. Das Tierreich 94:256, 258, 267 >> Neoperla
  • Zwick, P. 1973. Folia Entomologica Hungarica Rovartani Kozlemenyek (Series Nova) 26-Supplement:381-393 >> Neoperla
  • Zwick, P. 1973. Annales Zoologici 30(16):471-507 >> Note: Revision of types of Enderlein. >> Neoperla
  • Bednarik & McCafferty. 1977. The Great Lakes Entomologist 10(4):224   >> Note: as Neoperla clymene whose concept was not well set at the time. >> Neoperla
  • Stark & Baumann. 1978. Gr. Bas. Nat. 38(1):97-114   >> Neoperla
  • Zwick, P. 1981. Orient. Ins. 15(2):113-126 >> Neoperla
  • Zwick, P. 1983. Spixiana 6(2):167-204   >> Neoperla
  • Ernst & Stewart. 1985. Can. J. Zool. 63:2962-2968 >> Note: as N. clymene >> Neoperla
  • Zwick, P. 1986. Aquatic Insects 8(1):1-53 >> Neoperla
  • Ernst & Stewart. 1986. Aquatic Insects 8(4):237-254 >> Neoperla
  • Stark. 1987. Aquatic Insects 9(1):45-50 >> Neoperla
  • Harman. 1997. Journal of Freshwater Ecology 12(3):469   >> Neoperla
  • Du, Y-Z. & He. 2001. In Dominguez [Ed.]. Trends in Research in Ephemeroptera and Plecoptera. 369-375 >> Neoperla
  • Ogbogu. 2006. Illiesia 2(4):27-30   >> Neoperla
  • Harp & Robison. 2006. Journal of the Arkansas Academy of Science 60:55 >> Neoperla
  • Stark & Sivec. 2007. Illiesia 3(7):53-64   >> Neoperla
  • Stark & Sivec. 2008. Illiesia 4(3):52   >> Note: Key to males from Vietnam. >> Neoperla
  • Isobe, Yoshimura & Oishi. 2008. In Hauer, Stanford & Newell [Ed.]. International Advances in the Ecology, Zoogeography, and Systematics of Mayflies and Stoneflies 51-60   >> Neoperla
  • Stark & Sheldon. 2009. Illiesia 5(2):11   >> Neoperla
  • Lenat, Zuellig, Kondratieff & Beaty. 2009. Illiesia 5(15):164-168   >> Neoperla
  • Sivec & Stark. 2011. Illiesia 7(24):264-279   >> Neoperla
  • Harrison, A.B. & Morse. 2011. Trans. Amer. Ent. Soc. 138(1-2):39 >> Neoperla
  • Hamid & Rawi. 2011. Aquatic Insects 33(4):281-292 >> Neoperla
  • Sivaramakrishnan, Arunachalam, Sundar & Selva Kumar. 2011. In Tyagi & Veer. Entomology Ecology & Biodiversity 109 >> Neoperla
  • Li, W-H., Guoquan Wang, W. Li & Murányi. 2012. Zootaxa 3597:15-24 >> Neoperla
  • McPherson, Turner & Whiles. 2013. The Great Lakes Entomologist 46(1-2):42-89 >> Neoperla
  • Li, W-H., Guoquan Wang & Qin. 2013. ZooKeys 290:21-30   >> Neoperla
  • Li, W-H., H. Liang & W. Li. 2013. Zootaxa 3652(3):353-369   >> Neoperla
  • Qin, Murányi, Guoquan Wang & W-H. Li. 2013. ZooKeys 326:1-16   >> Neoperla
  • Li, W-H., Murányi & R. Wang. 2014. Zootaxa 3753(1):1-9 >> Neoperla
  • Murányi, W-H. Li, Jeon, J. M. Hwang & Seo. 2015. Zootaxa 3918(1):113-127 >> Neoperla
  • Stark & Sivec. 2015. Illiesia 11(7):75-91   >> Neoperla
  • Anzani, Krisanti & Wardiatno. 2016. Journal of Biodiversity and Environmental Sciences 9(1):128 >> Neoperla
  • Chen, Z-T. & Y-Z. Du. 2018. Zootaxa 4375(1):64   >> Neoperla
  • Yang, D. & W-H. Li. 2018. Species Catalogue of China. Vol. 2. Animals, Insecta (III), Plecoptera 36 >> Neoperla
  • Mo, R. Liu, W-H. Li & M. Shi. 2020. Zootaxa 4758(2):375 >> Note: Checklist of species known from China before 1999. >> Neoperla
  • Pelingen & Freitag. 2020. Zootaxa 954:58 >> Note: Checklist of Neoperla from the Philippines. >> Neoperla
• Type species:  Perla occipitalis Pictet, F.J., inherited from replaced name; authority:  Needham. 1905. Proc.Biol.Soc.Wash. 18:107-110
• The first family-group name based on this genus is Neoperlinae Enderlein, 1909.

• synonym Formosina Klapálek, 1913
• synonym Formosita Klapálek, 1914
• synonym Javanita Klapálek, 1909
• synonym Ochthopetina Enderlein, 1909
• synonym Oodeia Klapálek, 1921
• synonym Pseudoperla Banks, 1892
• synonym Simpliperla Wu, 1962
• synonym Tropidogynoplax Enderlein, 1910
• nomen dubium brachyura Navás, 1918
• nomen dubium corvicana Navás, 1927
• nomen dubium laminulata (Enderlein, 1909)
• nomen dubium pastellsi Navás, 1925
• nomen dubium tokiensis Navás, 1920
• nomen dubium vitalisi (Navás, 1923)
• nomen nudum caudalis
• species BoM [temporary name]
• species Id-A [temporary name]
• species Id-B [temporary name]
• species Id-C [temporary name]
• species Id-D [temporary name]
• species Vn-A [temporary name]
• species Vn-B [temporary name]
• species Vn-C [temporary name]
• species group clymene Zwick, 1983
• synonym aeripennis Zwick, 1983
• synonym clymene Zwick, 1983
• species aeripennis (Enderlein, 1909)
• species agumbe Stark & Sivec, 2015
• species aliqua Zwick, 1973
• species anjiensis Yang & Yang, 1998
• species asperata Zwick, 1988
• species baotianmana Li & Wang, 2012
• species breviscrotata Du, 1999
• species brevistyla Li & Murányi, 2013
• species caligata (Burmeister, 1839)
• species cavaleriei (Navás, 1922)
• species chebalinga Chen & Du, 2016
• species clara Stark & Sivec, 2008
• species clymene (Newman, 1839)
• species coronata Zwick, 1988
• species daklak Stark & Sivec, 2008
• species distincta Zwick, 1983
• species duratubulata Du, 1999
• species edmundsi Stark, 1983
• species emarginata Stark & Sivec, 2015
• species formosana Okamoto, 1912
• species foveolata Klapálek, 1921
• species gordonae Stark, 1983
• species hajeki Wang, Li & Li, 2022
• species hoabihnica Navás, 1932
• species hubeiensis Li & Wang, 2012
• species ignacsiveci Li & Li, 2013
• species infuscata Wu, 1935
• species jacobsoni Klapálek, 1910
• species laotica Zwick, 1988
• species leptacantha Stark & Sivec, 2008
• species leptophallus Zwick, 1988
• species lii Du, 1999
• species lui Du & Sivec, 2004
• species microtumida Wu & Claassen, 1934
• species monacha Stark & Sivec, 2008
• species nova Zwick, 1988
• species orissa Stark & Sivec, 2015
• species pani Chen & Du, 2016
• species parva Banks, 1939
• species primitiva Geijskes, 1952
• species propinqua Zwick, 1983
• species rigidipenis Zwick, 1983
• species rougemonti Zwick, 1986
• species sabah Zwick, 1986
• species saraburi Zwick, 1988
• species sarawak Zwick, 1986
• species sauteri Klapálek, 1912
• species serrata Zwick, 1988
• species sinuata Stark & Sivec, 2008
• species starki Zwick, 1986
• species sumatrana (Enderlein, 1909)
• species sungi Cao & Bae, 2007
• species theobromae Zwick, 1986
• species uniformis Banks, 1937
• species vesperi Zwick, 1983
• species group montivaga
• synonym diehli
• synonym luteola Zwick, 1983
• synonym montivaga
• species sp.1 [temporary name]
• species sp.2 [temporary name]
• species subgroup diehli Sivec, 1985
• species subgroup recta Banks, 1913
• species subgroup sinensis Chu, 1928
• species adamantea Murányi & Li, 2015
• species affinis Zwick, 1983
• species annulatispina Mo, Li & Wang, 2021
• species baisha Kong & Li, 2014
• species baishuijiangensis Du, 2005
• species banksi (Illies, 1966)
• species bicolor Yang, Zhang & Li, 2017
• species bicornuta Yang & Yang, 1995
• species bicurvata Kong & Li, 2014
• species bilobata Zwick, 1986
• species biprojecta Du, 2001
• species borneensis (Enderlein, 1909)
• species cameronis Zwick, 1988
• species chui Wu & Claassen, 1934
• species costalis (Klapálek, 1913)
• species curvispina Wu, 1948
• species dao Stark & Sivec, 2008
• species dashahena Du, 2005
• species dorsispina Yang & Yang, 1996
• species erecta Stark & Sivec, 2008
• species falcatata Mo, Wang, Li & Murányi, 2021
• species fallax Klapálek, 1910
• species fanjingshana Yang & Yang, 1992
• species flavescens Chu, 1929
• species flexiscrotata Du, 2000
• species furcata Zwick, 1986
• species furcifera Klapálek, 1909
• species furcomaculata Kong & Li, 2016
• species gaoligongshana Wang, Li & Li, 2022
• species goguryeo Murányi & Li, 2015
• species guizhouensis Yang & Yang, 1991
• species hainanensis Yang & Yang, 1995
• species hamata Jewett, 1975
• species harina Navás, 1929
• species henana Li, Wu & Zhang, 2011
• species hubbsi Ricker, 1952
• species hubleyi Stark & Sivec, 2008
• species idella Stark & Sivec, 2008
• species illiesi Zwick, 1983
• species jigongshana Li & Li, 2014
• species klapaleki Banks, 1937
• species latamaculata Du, 2005
• species latispina Wang & Li, 2013
• species leigongshana Du & Wang, 2007
• species lieftincki Zwick, 1983
• species lihuae Li & Murányi, 2014
• species limbatella Navás, 1933
• species longispina Wu, 1938
• species longwangshana Yang & Yang, 1998
• species lushana Wu, 1935
• species luteola (Burmeister, 1839)
• species magisterchoui Du, 2000
• species malleus Zwick, 1988
• species melanocephala Navás, 1931
• species mesospina Li & Wang, 2013
• species montivaga Zwick, 1977
• species multispinosa Stark & Sivec, 2008
• species naviculata Klapálek, 1909
• species nebulosa Stark & Sivec, 2008
• species oculata Banks, 1924
• species perspicillata Zwick, 1980
• species pilosella Klapálek, 1905
• species qingyuanensis Yang & Yang, 1995
• species qinlingensis Du, 2005
• species quadrata Wu & Claassen, 1934
• species rotunda Wu, 1948
• species schlitz Stark & Sivec, 2008
• species securifera Zwick, 1986
• species shangsiensis Mo, Wang, Li & Murányi, 2021
• species signatalis Banks, 1937
• species similidella Li & Wang, 2013
• species similiflavescens Li & Zhang, 2014
• species similiserecta Wang & Li, 2012
• species simplicior Navás, 1932
• species song Stark & Sivec, 2008
• species spinaloba Stark & Sivec, 2008
• species stueberae Zwick, 1983
• species tadpolata Li & Murányi, 2014
• species taibaina Du, 2005
• species taihorinensis (Klapálek, 1913)
• species taiwanica Sivec & Zwick, 1987
• species tamdao Cao & Bae, 2007
• species teresa Stark & Sivec, 2008
• species tetrapoda Zwick, 1986
• species thai Stark, 1983
• species tingwushanensis Wu, 1935
• species transversprojecta Du & Sivec, 2004
• species truncata Wu, 1948
• species tuberculata Wu, 1937
• species wui Yang & Yang, 1990
• species xuansongae Li & Li, 2013
• species yanlii Li & Wang, 2014
• species yaoshana Li, Wang & Lu, 2011
• species yentu Cao & Bae, 2007
• species yingshana Chen & Du, 2017
• species yunnana Wang, Murányi & Li, 2014
• species zonata Stark & Sivec, 2008
• species aestiva Uchida, 1990
• species africana Klapálek, 1909
• species agtouganon Sivec & Stark, 2011
• species agusani Sivec, 1984
• species alboguttata Zwick, 1986
• species angulata (Walker, 1852)
• species asperipenis Zwick, 1980
• species atropennis Banks, 1924
• species bicoronata Zwick, 1986
• species bimaculata Li, Wang & Li, 2021
• species biseriata Zwick & Anbalagan, 2007
• species bredoana Navás, 1932
• species caii Li, Wang & Wang, 2017
• species carlsoni Stark & Baumann, 1978
• species catharae Stark & Baumann, 1978
• species choctaw Stark & Baumann, 1978
• species connectens Zwick, 1986
• species coosa Smith & Stark, 1998
• species coralliata Uchida, 1990
• species coreensis Ra, Kim, Kang & Ham, 1994
• species coxi Stark, 1995
• species darlingi Stark & Sivec, 2007
• species dayak Zwick, 1986
• species dentata Sivec, 1984
• species divergens Zwick, 1986
• species dolichocephala Klapálek, 1909
• species emeishana Li, Wang & Wang, 2017
• species falayah Stark & Lentz, 1988
• species flavicincta Zwick, 1985
• species flinti Sivec, 1984
• species gaufini Stark & Baumann, 1978
• species geniculata (Pictet, 1841)
• species geniculatella Okamoto, 1912
• species grafei Stark & Sheldon, 2009
• species harperi Zwick, 1980
• species harpi Ernst & Stewart, 1986
• species harrisi Stark & Lentz, 1988
• species hatakeyamae Okamoto, 1912
• species hermosa Banks, 1924
• species indica Needham, 1909
• synonym indica Klapálek, 1909
• species inexspectata Zwick, 1980
• species inutilis Zwick, 1973
• synonym sumatrana Klapálek, 1909
• species jewetti Sivec, 1984
• species katmanduana Harper, 1977
• species kunenensis (Barnard, 1934)
• synonym kunensis Illies, 1966
• species lahu Stark, 1983
• species leroiana Klapálek, 1911
• species limushana Mo, Wang & Li, 2021
• species mainensis Banks, 1948
• species minuta Du & Chen, 2016
• species moesta Banks, 1939
• species multilobata Zwick, 1986
• species nigra Sivec, 1984
• species niponensis (McLachlan, 1875)
• species nitida Kimmins, 1950
• species obliqua Banks, 1913
• synonym apoana Banks, 1937
• species obscura Zwick, 1981
• species obscurofulva (Wu, 1962)
• species occipitalis (Pictet, 1841)
• synonym freytagi Stark & Baumann, 1978
• species ochracea Zwick, 1981
• species osage Stark & Lentz, 1988
• species palawan Sivec & Stark, 2011
• species pallescens Banks, 1937
• species pallicornis Banks, 1937
• species pangantihoni Pelingen & Freitag, 2021
• species paucispinosa Zwick, 1986
• species peterzwicki Stark & Sivec, 2008
• species philippina Sivec, 1984
• species pluvia Uchida, 1990
• species punan Zwick, 1986
• species ramosa (Navás, 1919)
• species reticulata Zwick, 1986
• species robisoni Poulton & Stewart, 1986
• species sabang Sivec & Stark, 2011
• species salakot Sivec & Stark, 2011
• species schmidi Aubert, 1959
• synonym complicata Sivec, 1980
• species schmidiana Zwick, 1981
• species separanda Zwick, 1983
• species seriespinosa Zwick, 1986
• species sexlobata Chen & Du, 2016
• species shiwandashana Mo, Yao, Wang & Li, 2019
• species silvaeae Zwick, 1986
• species sitahoanensis Sivec, 1985
• species siveci Zwick, 1980
• species spinosa Zwick, 1986
• species spio (Newman, 1839)
• synonym annulata (Navás, 1923)
• synonym arambourgana Navás, 1931
• synonym bottegoana Navás, 1933
• synonym burgeoni Navás, 1926
• synonym camerunensis Enderlein, 1909
• synonym conradti (Enderlein, 1909)
• synonym didita (Enderlein, 1909)
• synonym dubia Klapálek, 1909
• synonym excisa Klapálek, 1909
• synonym haugi (Navás, 1916)
• synonym innexa (Navás, 1920)
• synonym lacroixi Lestage, 1921
• synonym laticollis Klapálek, 1909
• synonym leopoldina Navás, 1932
• synonym luluana Navás, 1931
• synonym needhami Lestage, 1921
• synonym nigricauda Klapálek, 1909
• synonym petiolata (Navás, 1920)
• synonym seyrigiana Navás, 1934
• synonym sjöstedti Klapálek, 1909
• synonym tenera (Navás, 1915)
• synonym transvaalensis (Enderlein, 1909)
• synonym lujana Navás, 1931
• species stewarti Stark & Baumann, 1978
• species tenuispina Klapálek, 1921
• species tibetensis Li & Liu, 2019
• species tortipenis Zwick, 1980
• species triangulata Kawai, 1975
• species unicolor Zwick, 1986
• species ussurica Sivec & Zhiltzova, 1996
• synonym ussurica Sivec & Zhiltzova, 1995
• species vallis Uchida, 1990
• species variegata Klapálek, 1909
• synonym centralis Banks, 1939
• synonym lebangina Navás, 1929
• species venosa Kimmins, 1950
• species verna Uchida, 1990
• species wagneri Sivec, 1984
• species wuzhishana Chen & Du, 2016
• species zhiltzovae Teslenko, 2012

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